I was able to obtain a B&W print of this paper from my local library in MN. Contrary to the initial post, and also contrary to what I could get from the abstract, all named taxa of subgenus
Yaotriton were included for the first time, but not all members of genus
Tylototriton. This includes confirmation of species status for
T.notialis as well as a [known] range extension to Vietnam.
T.hainanensis is not yet confirmed outside of Hainan, but the species list for Vietnam now includes
T.notialis, T.ziegleri, T.asperrimus, and
T.vietnamensis. Subgenus
Tylototriton likely also occurs in Vietnam (probably more than one species) but requires confirmation. While no status change is proposed, the differences between
dabienicus and
wenxianensis are similar to those between other full species, and I continue to treat these two as species. Because of the very few species and localitites of subgenus
Tylototriton included, it's not clear whether the three species included are accurately identified.
All members of the subgenus are distinguished by morphological traits. Among those for
T.ziegleri are well-developed cranial ridges, which are unique for the subgenus. The skin is roughest, the ridge most tubercular and segmented, and rib nodules most distinct within the subgenus. Coloration is mainly unreliable as a distinguishing trait.
Taxa included:
T.asperrimus
T.broadoridgus
T.dabienicus
T.hainanensis
T.lizhenchangi
T.notialis
T.vietnamensis
T.wenxianensis
No additional unnamed species are included, which does not preclude their existence.
Diagnosis, quoted in full from the paper:
A medium-sized newt of the genus Tylototriton, SVL 54.4-68.3 mm in males and 70.8mm in the female; skin rough in fine granules; bony ridges on head distinct; vertebral ridge prominent and segmented, forming a row of tubercles; rib nodules prominent; limbs long and thin; tips of forelimbs and hindlimbs greatly overlapping when adpressed along body; tail thin; dorsum uniform dark brown or blackish; rib nodules, finger and toe tips, parts of soles and palms, and vent continuing to ventral ridge of tail bright orange.