I also have a coupolke like this and a couple that are very dark (no oranbge knobbles). I have seen light colors like this in alpestris too , for instance when they go into water with a muddy clay bottom substrate. In this way they mix in better with the bottom substrate.
The new Vietnamese tylototritons also have that : on land they are black in the water they get lighter. Maybe try to set them on a lighter bottomground might return those colors ??
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