Thanks for the replies! I wasn't convinced it is asperrimus due to the lack or less defined knobbles compared to this individual taken near the type locality
A good conclusion, except that the link provided is for an animal which is not from near the type locality of
T.asperrimus (in southern Guizhou China), but rather from Ha Giang province, Vietnam. It may or may not be that species, and is geographically much closer to Cao Bang animals which are known to be an undescribed species currently called
T.cf.vietnamensis.
My first thought is that at least the animal in the center looks an awful lot like
T.taliangensis. Detailed photos of the head, shoulders, and front torso; and of the underside including cloaca and tail; would be helpful.
T.shanjing, T.cf.shanjing, T.kweichowensis, T.cf.kweichowensis, and
T.notialis are all ruled out easily by coloration. All of these species have brightly pigmented costal warts.
T.verrucosus and
T.vietnamensis are ruled out by their non-black dorsal colors and distinct costal warts [flattened in the latter].
T.asperrimus is ruled out by its prominent costal warts.
That leaves
T.taliangensis,
T.wenxianensis, T.dabienicus, T.hainanensis, and
T.cf.vietnamensis.
T.cf.vietnamensis apparently has prominent costal warts and can be ruled out.
T.dabienicus is thus far only known from a protected forest region and is perhaps unlikely to be collected...so far. i wouldn't rule it out, but consider it unlikely.
T.hainanensis has distinct, flattened costal warts.
T.wenxianensis [and
T.dabienicus] have indistinct costal warts which may form a ridge.
T.taliangensis is the only dorsally black species with red parotids, and has indistinct costal warts.
I think that reduces it
T.wenxianensis and
T.taliangensis, and it looks like you have both. The two species occur relatively closely together and stand a good chance of being shipped in mixed batches. The all-black ones might be
T.hainanensis, but without more clear images of the warts, I suspect not.
I've updated my map, linked in the
T.verrucosus ID thread. I've changed some Boehme et al records of
T.vietnamensis, which are treated as
T.cf.vietnamensis by Stuart et al. I've also deleted an incorrect locality of
T.asperrimus [error in longitude was mine] and two records which were localities that now have more precise information and updated taxonomy. Those changes combined make all known taxa of
Tylototriton geographically consistant. The previous map had overlapping ranges for vietnamensis, asperrimus, and cf. vietnamensis.
I am still seeking out documented localities for Sikkim, Arunachal Pradesh, Meghalaya, Assam, and Bhutan. There should be localities in Tibet, Bihar, and Bangladesh as well, but I know of no records. [all
T.verrucosus] I am also lacking most records for China. Almost every province east of southern Gansu, Sichuan, and Yunnan should have records, though few do and I lack data on most of those.